The University of Western Australia

UWA Staff Profile

Ipsum Lorem

Assoc/Prof Joseph Tomkins

Associate Professor
Biological Sciences, School of

Contact details
Address
School of Biological Sciences
The University of Western Australia (M092)
35 Stirling Highway
CRAWLEY WA 6009
Australia
Phone
+61 8 6488 2110
Email
joseph.tomkins@uwa.edu.au
Personal homepage
http://alternativetactics.com/
Location
Room 2.15, Zoology Building, Perth campus

Senior Research Fellow/Future Fellow
Evolutionary Biology, Centre for

Contact details
Address
Centre for Evolutionary Biology
The University of Western Australia (M092)
35 Stirling Highway
CRAWLEY WA 6009
Australia
Phone
+61 8 6488 2110
Email
joseph.tomkins@uwa.edu.au
Personal homepage
http://alternativetactics.org/
Location
Room 2.15, Zoology Building, Perth campus
Qualifications
BSc Sheff., PhD Liv.
Publications
Tomkins, J. L., Kotiaho, J. S. & LeBas, N. R. 2005 Maters of scale: positive allometry and the evolution of male dimorphisms. American Naturalist. 165, 389-402.



Rutstein, A. N., Gilbert, L. & Tomkins, J. L. 2005 Experience counts: lessons from studies of differential allocation. Behavioral Ecology. 16, 957-960.



Tomkins, J. L., Kotiaho, J. S. & LeBas, N. R. 2005 Phenotypic plasticity in the developmental integration of morphological trade-offs and secondary sexual trait compensation. Proceedings of the Royal Society of London B. 273, 543-551.



Tomkins, J. L. & Brown, G. S. 2004 Population density drives the local evolution of a threshold dimorphism. Nature. 431, 1099-1103.



Tomkins, J. L., Radwan, J., Kotiaho, J. S. & Tregenza, T. 2004 Genic capture and resolving the lek paradox. Trends in Ecology and Evolution 19, 323-328.



Tomkins, J. L., LeBas, N. R., Unrug, J. & Radwan, J. 2004. Testing the status-dependent ESS: population variation in fighter expression in the mite Sancassania berlesei. Journal of Evolutionary Biology 17, 1377-1388.



Knell, R. K., Pomfret, J. C. & Tomkins, J. L. 2004 The limits of elaboration: curved allometries reveal the constraints on mandible size in stag beetles. Proceedings of the Royal Society of London B 271, 523-528.



Unrug, J., Tomkins, J. L. & Radwan, J. 2004 Alternative phenotypes and sexual selection: can dichotomous handicaps honestly signal quality? Proceedings of the Royal Society of London B 271, 1401-1406.



Tomkins, J. L. & Kotiaho, J. S. 2004 Publication bias in meta-analysis: seeing the wood for the trees. Oikos 104, 194-196.



Tomkins, J. L. & Simmons, L. W. 2003 Fluctuating asymmetry and sexual selection: paradigm shifts, publication bias and observer expectation. In Developmental Instability: Causes and Consequences (ed. M. Polak), pp. 231-261. New York: Oxford University Press.



Kotiaho, J. S., Simmons, L. W., Hunt, J. & Tomkins, J. L. 2003 Males influence maternal effects that promote sexual selection. American Naturalist 161, 852-859.



Tomkins, J. L. & Simmons, L. W. 2002 Measuring relative investment: a case study of testes investment in species with alternative male reproductive tactics. Animal Behaviour 63, 1009-1016.



Kotiaho, J. S. & Tomkins, J. L. 2002 Meta-analysis can it ever fail ? Oikos 96, 551-553.



Radwan, J., Unrug, J. & Tomkins, J. L. 2002 Status dependence and morphological trade-offs in the expression of a sexually selected character in the mite, Sancassania berlesei. Journal of Evolutionary Biology 15, 744-752.



Colegrave, N., Kotiaho, J. S. & Tomkins, J. L. 2002 Mate choice or polyandry: reconciling genetic compatibility and good genes sexual selection. Evolutionary Ecology Research 2002, 911-917.



Kotiaho, J. S. & Tomkins, J. L. 2001 The discrimination of alternative male morphologies. Behavioral Ecology 12, 553-557.



Kotiaho, J. S., Simmons, L. W. & Tomkins, J. L. 2001 Towards a resolution of the lek paradox. Nature 410, 684-686.



Tomkins, J. L., Simmons, L. W. & Alcock, J. 2001 Brood provisioning strategies in Dawsons burrowing bee, Amegilla dawsoni (Hymenoptera: Anthophorini). Behavioral Ecology and Sociobiology 50, 81-89.



Tomkins, J. L. & Kotiaho, J. S. 2001 Fluctuating asymmetry. In Encyclopedia of Life Sciences. London: MacMillan Reference Ltd.



Simmons, L. W., Tomkins, J. L. & Alcock, J. A. 2000 Can minor males of Dawson's burrowing bee, Amegilla dawsoni (Hymenoptera: Anthophorini) compensate for reduced access to virgin females through sperm competition ? Behavioral Ecology 11, 319-325.



Tomkins, J. L. & Simmons, L. W. 2000 Sperm competition games played by dimorphic male beetles: fertilisation gains with equal mating success. Proceedings of the Royal Society of London B 267, 1547-1553.



Tomkins, J. L. & Simmons, L. W. 1999 Heritability of size but not symmetry in a sexually selected trait chosen by female earwigs. Heredity 82, 151-157



Tomkins, J. L. 1999 The ontogeny of asymmetry in earwig forceps. Evolution 53, 157-163.





Simmons, L. W., Tomkins, J. L., Kotiaho, J. S. & Hunt, J. 1999 Fluctuating paradigm. Proceedings of the Royal Society of London B 266, 593-595.



Tomkins, J. L. 1999 Environmental and genetic determinants of the male forceps length dimorphism in the European earwig Forficula auricularia. L. Behavioral Ecology and Sociobiology 47, 1-8.



Simmons, L. W., Tomkins, J. L. & Hunt, J. C. 1999 Sperm competition games played by dimorphic male beetles. Proceedings of the Royal Society of London B 266, 145-150.



Kotiaho, J., S., Tomkins, J. L. & Simmons, L. W. 1999 Unfamiliar citations breed mistakes. Nature 400, 307.



Hunt, J., Kotiaho, J. S. & Tomkins, J. L. 1999 Dung pad residence time covaries with male morphology in the dung beetle Onthophagus taurus. Ecological Entomology 24, 174-180.



Tomkins, J. L., Simmons, L. W., Knell, R. J. & Norris, K. A. 1999 Correlates of ball size and rolling speed in the dung beetle Kheper nigroaeneus (Coleoptera : Scarabaeidae). Journal of Zoology 248, 483-487.



Tomkins, J. L. & Simmons, L. W. 1998 Female choice and manipulations of forceps size and symmetry in the European earwig Forficula auricularia L. Animal Behaviour 56, 347-356.



Tomkins, J. L. 1997 Sexual Selection and the Evolution of Earwig Forceps: The University of Liverpool.



Simmons, L. W. & Tomkins, J. L. 1996 Sexual selection and the allometry of earwig forceps. Evolutionary Ecology 10, 97-104.



Tomkins, J. L. & Simmons, L. W. 1996 Dimorphisms and fluctuating asymmetry in the forceps of male earwigs. Journal of Evolutionary Biology. 9, 753-770.



Tomkins, J. L. & Simmons, L. W. 1995 Patterns of fluctuating asymmetry in earwig forceps: no evidence for reliable signalling. Proceedings of the Royal Society of London B. 259, 89-96.



Simmons, L. W., Tomkins, J. L. & Manning, J. T. 1995 Sampling bias and fluctuating asymmetry. Animal Behaviour 49, 1697-1699.

Research profile
Research profile and publications
 

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Tuesday, 3 November, 2015 2:39 PM